Senna (plant)

 

Senna (plant)

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Senna
Cassia senna Ypey80-cropped.jpg
Senna alexandrina
Scientific classificatione
Kingdom:Plantae
Clade:Tracheophytes
Clade:Angiosperms
Clade:Eudicots
Clade:Rosids
Order:Fabales
Family:Fabaceae
Tribe:Cassieae
Genus:Senna
Mill.
Type species
Senna alexandrina
Mill.
Species

Over 300; see text

Synonyms[1]

Senna, the sennas, is a large genus of flowering plants in the legume family (Fabaceae, subfamily Caesalpinioideae,[2] tribe Cassieae). This diverse genus is native throughout the tropics, with a small number of species in temperate regions. The number of species is estimated to be from about 260[3] to 350.[4] The type species for the genus is Senna alexandrina. About 50 species of Senna are known in cultivation.[5]

Description

Senna includes herbs, shrubs, and trees. The leaves are pinnate with opposite paired leaflets. The inflorescences are racemes at the ends of branches or emerging from the leaf axils. The flower has five sepals and five usually yellow petals. There are ten straight stamens. The stamens may be different sizes, and some are staminodes. The fruit is a legume pod containing several seeds.[6]

Systematics

Chamaecrista, Cassia, and Senna form a monophyletic group which some authors have called Cassia sensu lato.[7] In 1982, the group was named Cassiinae and classified as a subtribe of the tribe Cassieae.[8] The tribe Cassieae contains 21 genera and is now known to be polyphyletic,[7] but the classification is still accepted because a revision of Fabaceae has yet to be published.[7]

The genus Senna has had a complex taxonomic history.[9] What is now known as Senna was included by Linnaeus in his concept of Cassia in Species Plantarum in 1753.[10] Philip Miller segregated Senna from Cassia in 1754 in the fourth edition of The Gardeners Dictionary.[11] Until 1982, many authors, following Linnaeus, did not recognize Senna and Chamaecrista, but included them in a broadly circumscribed Cassia sensu lato. Phylogenetic analyses of DNA have shown that Chamaecrista, Cassia, and Senna are all monophyletic, but the relationships between these three genera have not been resolved.[3] They are therefore shown in phylogenetic trees as a tritomy.

Etymology

The genus name derives from the Arabic sanā, describing plants whose leaves and pods have catharitic and laxative properties.[12]

Ecology

The caterpillars of many Lepidoptera species feed on Senna plants. The black witch (Ascalapha odorata), two-barred flasher (Astraptes fulgerator), common emigrant (Catopsilia pomona), and mottled emigrant (C. pyranthe) have all been recorded on candle bush (S. alata), for example.

Pollination

Senna species are pollinated by a variety of bees, especially large female bees in genera such as Xylocopa.[3] They rely on "buzz pollination" and some within that on "ricochet pollination", which is a secondary pollen presentation where the pollen is not deposited on the pollinator's body by direct contact with the anthers.[13] The flowers have two sets of stamen: feeding stamens, which are longer, and pollinating stamens, which are smaller in size.[14] Due to buzz pollination, the pollens from the pollinating stamens get thrown from the anthers and ricochets against the petals multiple times before it settles on the dorsal side of the pollinating bee. The roughness on the petal walls causes the pollen to slow down its speed. The ricocheting effect alone cannot ensure effective pollen dissemination. It is aided by static charges wherein the flying bees become positively charged owing to the friction in the air and the pollen becomes negatively charged because of which they naturally get attracted to the bees body. The pollinator bee ends up carrying the pollen and also gets to feed on the pollen which is on the feeding stamens.[13] Some species also have extrafloral nectaries on the leaves or flower stalks, which attract ants, but do not benefit pollinators.[3]

Uses

Some Senna species are used as ornamental plants in landscaping. The species is adapted to many climate types.

Cassia gum, an extract of the seeds of Chinese senna (S. obtusifolia), is used as a thickening agent. The leaves and flowers of Siamese cassia (S. siamea) are used in some Southeast Asian cuisines, such as Thai, Shan/Burmese and Lao cuisines. They are known as khi-lek in Thai, and are used in curries.[15]

Laxative

Historically, Egyptian senna (S. alexandrina) was used as a laxative in the form of senna pods, or as herbal tea made from the leaves.[citation needed] Senna is considered to be a bowel stimulant on the myenteric plexus of the colon to induce peristaltic contractions and decrease water absorption from inside the colon, effects that would provide relief from constipation.[16]

Senna or its extracted sennosides, alone or in combination with sorbitol or lactulose, have been evaluated in systematic reviews and Cochrane reviews for treatment of constipation in children and the elderly. Some studies showed limited evidence for efficacy,[17][18][19] whereas others indicated the study designs were too weak to be certain of senna having utility as a laxative.[16][20][21]

Species

As of December 2019, Plants of the World Online accepts the following species:

Senna alexandrina
Senna artemisioides
Senna bicapsularis
Senna didymobotrya
Senna hirsuta
Senna lindheimeriana
Senna macranthera
Senna multiglandulosa
Senna occidentalis
Senna pendula
Senna pinheiroi
Senna spectabilis
Senna surattensis
  • Senna acanthoclada (Griseb.) H.S.Irwin & Barneby
  • Senna acclinis (F.Muell.) Randell
  • Senna aciphylla (Benth. ex A.Gray) Randell
  • Senna aculeata (Pohl ex Benth.) H.S.Irwin & Barneby
  • Senna acunae (Borhidi) A.Barreto & Yakovlev
  • Senna acuparata H.S.Irwin & Barneby
  • Senna acuruensis (Benth.) H.S.Irwin & Barneby
  • Senna acutisepala (Benth.) H.S.Irwin & Barneby
  • Senna affinis (Benth.) H.S.Irwin & Barneby
  • Senna alata (L.) Roxb. – candlebush, Christmas candle
  • Senna alexandrina Mill. – Alexandrian senna
  • Senna andrieuxii (Benth.) H.S.Irwin & Barneby
  • Senna angulata (Vogel) H.S.Irwin & Barneby
  • Senna angustisiliqua (Lam.) H.S.Irwin & Barneby
  • Senna ankaranensis Du Puy & R.Rabev.
  • Senna anthoxantha (Capuron) Du Puy
  • Senna aphylla (Cav.) H.S.Irwin & Barneby
  • Senna apiculata (M.Martens & Galeotti) H.S.Irwin & Barneby
  • Senna appendiculata (Vogel) Wiersema
  • Senna apsidoneura (H.S.Irwin & Barneby) H.S.Irwin & Barneby
  • Senna araucarietorum H.S.Irwin & Barneby
  • Senna argentea (Kunth) H.S.Irwin & Barneby
  • Senna arida (Rose) H.S.Irwin & Barneby
  • Senna aristeguietae H.S.Irwin & Barneby
  • Senna armata (S.Watson) H.S.Irwin & Barneby
  • Senna arnottiana (Hook.) H.S.Irwin & Barneby
  • Senna artemisioides (Gaudich. ex DC.) Randell – grey cassia, leafless cassia, woody cassia
  • Senna atomaria (L.) H.S.Irwin & Barneby
  • Senna aurantia (Ruiz & Pav. ex G.Don) H.S.Irwin & Barneby
  • Senna auriculata (L.) Roxb. – avaram, Matara-tea, tanner's cassia
  • Senna aversiflora (Herb.) H.S.Irwin & Barneby
  • Senna aymara H.S.Irwin & Barneby
  • Senna baccarinii (Chiov.) Lock
  • Senna bacillaris (L.f.) H.S.Irwin & Barneby – West Indian showertree
  • Senna barclayana (Sweet) Randell
  • Senna barnebyana Lass. – pepper-leaf senna, smooth senna, yellow peabush
  • Senna barronfieldii (Colla) Hewson
  • Senna bauhinioides (A.Gray) H.S.Irwin & Barneby
  • Senna benitoensis (Britton & P.Wilson) H.S.Irwin & Barneby
  • Senna bicapsularis (L.) Roxb. – rambling senna, Christmas bush, money bush, yellow candlewood
  • Senna biglandularis A.O.Araujo & V.C.Souza
  • Senna birostris (Dombey ex Vogel) H.S.Irwin & Barneby
  • Senna bosseri Du Puy & R.Rabev.
  • Senna bracteosa D.B.O.S.Cardoso & L.P.Queiroz
  • Senna brongniartii (Gaudich.) H.S.Irwin & Barneby
  • Senna burkartiana (Villa) H.S.Irwin & Barneby
  • Senna cajamarcae H.S.Irwin & Barneby
  • Senna cana (Nees & Mart.) H.S.Irwin & Barneby
  • Senna candolleana (Vogel) H.S.Irwin & Barneby
  • Senna cardiosperma (F.Muell.) Randell
  • Senna catingae (Harms) L.P.Queiroz
  • Senna caudata (Standl.) H.S.Irwin & Barneby
  • Senna cearensis Afr.Fern.
  • Senna centranthera H.S.Irwin & Barneby
  • Senna cernua (Balb.) H.S.Irwin & Barneby
  • Senna chapmanii (Isely) A.Barreto & Yakovlev
  • Senna charlesiana (Symon) Randell
  • Senna chloroclada (Harms) H.S.Irwin & Barneby
  • Senna chrysocarpa (Desv.) H.S.Irwin & Barneby
  • Senna cladophylla (W.Fitzg.) Randell
  • Senna clavigera (Domin) Randell
  • Senna cobanensis (Britton) H.S.Irwin & Barneby
  • Senna coimbrae M.Nee & Barneby
  • Senna collicola H.S.Irwin & Barneby
  • Senna confinis (Greene) H.S.Irwin & Barneby
  • Senna corifolia (Benth.) H.S.Irwin & Barneby
  • Senna cornigera H.S.Irwin & Barneby
  • Senna coronilloides (Benth.) Randell
  • Senna corymbosa (Lam.) H.S.Irwin & Barneby – Argentine senna, Argentine wild sensitive plant[25]
  • Senna costata (J.F.Bailey & C.T.White) Randell
  • Senna covesii (A.Gray) H.S.Irwin & Barneby – Coues' cassia, Coves' cassia, desert senna[25]
  • Senna crassiramea (Benth.) H.S.Irwin & Barneby
  • Senna crotalarioides (Kunth) H.S.Irwin & Barneby
  • Senna cruckshanksii (Hook.) H.S.Irwin & Barneby
  • Senna cuatrecasasii H.S.Irwin & Barneby
  • Senna cumingii (Hook. & Arn.) H.S.Irwin & Barneby
  • Senna curvistyla (J.M.Black) Randell
  • Senna cushina (J.F.Macbr.) H.S.Irwin & Barneby
  • Senna cuthbertsonii (F.Muell.) Randell
  • Senna dardanoi Afr.Fern. & P.Bezerra
  • Senna dariensis (Britton & Rose) H.S.Irwin & Barneby
  • Senna davidsonii (V.Singh) V.Singh
  • Senna demissa (Rose) H.S.Irwin & Barneby
  • Senna didymobotrya (Fresen.) H.S.Irwin & Barneby – candelabra-tree, peanut butter cassia, African senna
  • Senna divaricata (Nees & Blume) Lock
  • Senna domingensis (Spreng.) H.S.Irwin & Barneby
  • Senna durangensis (Rose) H.S.Irwin & Barneby
  • Senna ellisiae (Brenan) Lock
  • Senna ferraria (Symon) Randell
  • Senna flexuosa (Randell) Randell
  • Senna × floribunda (Cav.) H.S.Irwin & Barneby
  • Senna foetidissima (Ruiz & Pav. ex G.Don) H.S.Irwin & Barneby
  • Senna formosa H.S.Irwin & Barneby
  • Senna fruticosa (Mill.) H.S.Irwin & Barneby
  • Senna galeottiana (M.Martens) H.S.Irwin & Barneby
  • Senna gardneri (Benth.) H.S.Irwin & Barneby
  • Senna garrettiana (Craib) H.S.Irwin & Barneby
  • Senna gaudichaudii (Hook. & Arn.) H.S.Irwin & Barneby – climbing cassia, heuhiuhi (Pacific Islands, Queensland)[26]
  • Senna georgica H.S.Irwin & Barneby
  • Senna glanduligera (H.St.John) A.C.Sm.
  • Senna glaucifolia (Randell) Randell
  • Senna glutinosa (DC.) Randell
  • Senna goniodes (A.Cunn. ex Benth.) Randell
  • Senna gossweileri (Baker f.) Lock
  • Senna guatemalensis (Donn.Sm.) H.S.Irwin & Barneby
  • Senna gundlachii (Urb.) H.S.Irwin & Barneby
  • Senna hamersleyensis (Symon) Randell
  • Senna harleyi H.S.Irwin & Barneby
  • Senna haughtii (J.F.Macbr.) H.S.Irwin & Barneby
  • Senna hayesiana (Britton & Rose) H.S.Irwin & Barneby
  • Senna hebecarpa (Fernald) H.S.Irwin & Barneby – American senna, wild senna
  • Senna heptanthera Randell
  • Senna herzogii (Harms) H.S.Irwin & Barneby
  • Senna hilariana (Benth.) H.S.Irwin & Barneby
  • Senna hirsuta (L.) H.S.Irwin & Barneby – woolly senna
  • Senna holosericea (Fresen.) Greuter
  • Senna holwayana (Rose) H.S.Irwin & Barneby
  • Senna hookeriana Batke
  • Senna huancabambae (Harms) H.S.Irwin & Barneby
  • Senna huidobriana (Phil.) Zoellner & San Martin
  • Senna huilana (Britton & Rose) H.S.Irwin & Barneby
  • Senna humifusa (Brenan) Lock
  • Senna incarnata (Benth.) H.S.Irwin & Barneby
  • Senna insularis (Britton & Rose) H.S.Irwin & Barneby
  • Senna intermedia (B.D.Sharma, Vivek. & Rathakr.) V.Singh
  • Senna italica Mill. – Port Royal senna, dog senna, Italian senna, Spanish senna
  • Senna itatiaiae H.S.Irwin & Barneby
  • Senna koelziana H.S.Irwin & Barneby
  • Senna kurtzii (Harms) H.S.Irwin & Barneby
  • Senna lactea (Vatke) Du Puy
  • Senna lasseigniana H.S.Irwin & Barneby
  • Senna latifolia (G.Mey.) H.S.Irwin & Barneby
  • Senna leandrii (Ghesq.) Du Puy
  • Senna lechriosperma H.S.Irwin & Barneby
  • Senna leiophylla (Vogel) H.S.Irwin & Barneby
  • Senna leptoclada (Benth.) Randell
  • Senna ligustrina (L.) H.S.Irwin & Barneby – privet senna
  • Senna lindheimeriana (Scheele) H.S.Irwin & Barneby – velvet-leaf senna, showy senna
  • Senna longiglandulosa (Benth.) H.S.Irwin & Barneby
  • Senna longiracemosa (Vatke) Lock
  • Senna loretensis (Killip) H.S.Irwin & Barneby
  • Senna lourteigiana H.S.Irwin & Barneby
  • Senna macranthera (DC. ex Collad.) H.S.Irwin & Barneby
  • Senna macrophylla (Kunth) H.S.Irwin & Barneby
  • Senna magnifolia (F.Muell.) Randell
  • Senna malaspinae H.S.Irwin & Barneby
  • Senna mandonii (Benth.) H.S.Irwin & Barneby
  • Senna manicula (Symon) Randell
  • Senna marilandica (L.) Link
  • Senna martiana (Benth.) H.S.Irwin & Barneby
  • Senna mensicola (H.S.Irwin & Barneby) H.S.Irwin & Barneby
  • Senna meridionalis (R.Vig.) Du Puy – Madagascar senna
  • Senna mexicana (Jacq.) H.S.Irwin & Barneby – Mexican senna
  • Senna mollissima (Humb. & Bonpl. ex Willd.) H.S.Irwin & Barneby
  • Senna monilifera H.S.Irwin & Barneby
  • Senna monozyx (H.S.Irwin & Barneby) H.S.Irwin & Barneby
  • Senna montana (B.Heyne ex Roth) V.Singh
  • Senna morongii (Britton) H.S.Irwin & Barneby
  • Senna mucronifera (Mart. ex Benth.) H.S.Irwin & Barneby
  • Senna multifoliolata (Paul G.Wilson) H.S.Irwin & Barneby
  • Senna multiglandulosa (Jacq.) H.S.Irwin & Barneby – glandular senna, downy senna
  • Senna multijuga (Rich.) H.S.Irwin & Barneby – false sicklepod
  • Senna mutisiana (Kunth) H.S.Irwin & Barneby
  • Senna nana (Benth.) H.S.Irwin & Barneby
  • Senna neglecta (Vogel) H.S.Irwin & Barneby
  • Senna nicaraguensis (Benth.) H.S.Irwin & Barneby
  • Senna nitida (Rich.) H.S.Irwin & Barneby
  • Senna notabilis (F.Muell.) Randell
  • Senna nudicaulis (Burkart) H.S.Irwin & Barneby
  • Senna obliqua (G.Don) H.S.Irwin & Barneby
  • Senna oblongifolia (Vogel) H.S.Irwin & Barneby
  • Senna obtusifolia (L.) H.S.Irwin & Barneby – coffee-weed, java-bean, American sicklepod
  • Senna occidentalis (L.) Link – antbush, coffee-senna, negro-coffee, septicweed, stinkingweed[27]
  • Senna oligoclada (F.Muell.) Randell
  • Senna orcuttii (Britton & Rose) H.S.Irwin & Barneby – Orcutt's senna
  • Senna organensis (Glaz. ex Harms) H.S.Irwin & Barneby
  • Senna oxyphylla (Kunth) H.S.Irwin & Barneby
  • Senna pachyrrhiza (L.Bravo) H.S.Irwin & Barneby
  • Senna pallida (Vahl) H.S.Irwin & Barneby
  • Senna papillosa (Britton & Rose) H.S.Irwin & Barneby
  • Senna paposana (Phil.) Zoellner & San Martin
  • Senna paradictyon (Vogel) H.S.Irwin & Barneby
  • Senna paraensis (Ducke) H.S.Irwin & Barneby
  • Senna pendula (Humb. & Bonpl. ex Willd.) H.S.Irwin & Barneby – Easter cassia
  • Senna pentagonia (Mill.) H.S.Irwin & Barneby
  • Senna peralteana (Kunth) H.S.Irwin & Barneby
  • Senna perrieri (R.Vig.) Du Puy
  • Senna petersiana (Bolle) Lock
  • Senna phlebadenia H.S.Irwin & Barneby
  • Senna phyllodinea (R.Br.) Symon
  • Senna pilifera (Vogel) H.S.Irwin & Barneby
  • Senna pilocarina (Symon) Randell
  • Senna pilosior (B.L.Rob.) H.S.Irwin & Barneby — Trans-Pecos senna
  • Senna pinheiroi H.S.Irwin & Barneby
  • Senna pistaciifolia (Kunth) H.S.Irwin & Barneby
  • Senna planitiicola (Domin) Randell
  • Senna pleurocarpa (F.Muell.) Randell – stripe-pod cassia[28]
  • Senna pneumatica H.S.Irwin & Barneby
  • Senna podocarpa (Guill. & Perr.) Lock
  • Senna polyantha (Collad.) H.S.Irwin & Barneby
  • Senna polyphylla (Jacq.) H.S.Irwin & Barneby
  • Senna praeterita H.S.Irwin & Barneby
  • Senna procumbens Randell
  • Senna pumilio (A.Gray) H.S.Irwin & Barneby – dwarf senna
  • Senna punoensis Lass.
  • Senna purpusii (Brandegee) H.S.Irwin & Barneby
  • Senna quinquangulata (Rich.) H.S.Irwin & Barneby
  • Senna racemosa (Mill.) H.S.Irwin & Barneby – limestone senna
  • Senna reniformis (G.Don) H.S.Irwin & Barneby
  • Senna reticulata (Willd.) H.S.Irwin & Barneby
  • Senna rigidicaulis (Burkart ex L.Bravo) H.S.Irwin & Barneby
  • Senna ripleyana (H.S.Irwin & Barneby) H.S.Irwin & Barneby – Ripley's senna
  • Senna rizzinii H.S.Irwin & Barneby
  • Senna robiniifolia (Benth.) H.S.Irwin & Barneby
  • Senna roemeriana (Scheele) H.S.Irwin & Barneby – twoleaf senna, Roemer senna
  • Senna rostrata (Mart.) H.S.Irwin & Barneby – New Mexico wild sensitive plant
  • Senna rugosa (G.Don) H.S.Irwin & Barneby
  • Senna ruiziana (G.Don) H.S.Irwin & Barneby
  • Senna rupununiensis H.S.Irwin & Barneby
  • Senna ruspolii (Chiov.) Lock
  • Senna saeri (Pittier) H.S.Irwin & Barneby
  • Senna sandwithiana H.S.Irwin & Barneby
  • Senna santanderensis (Britton & Killip) H.S.Irwin & Barneby
  • Senna scabriuscula (Vogel) H.S.Irwin & Barneby
  • Senna scandens (G.Don) H.S.Irwin & Barneby
  • Senna septemtrionalis (Viv.) H.S.Irwin & Barneby – arsenic-bush, Dooleyweed, laburnum, smooth senna
  • Senna sericea (Symon) Albr. & Symon
  • Senna siamea (Lam.) H.S.Irwin & Barneby – kassodtree, Siamese cassia, Thai cassia, Thailand shower
  • Senna silvestris (Vell.) H.S.Irwin & Barneby
  • Senna singueana (Delile) Lock
  • Senna skinneri (Benth.) H.S.Irwin & Barneby
  • Senna smithiana (Britton & Rose) H.S.Irwin & Barneby
  • Senna sophera (L.) Roxb.
  • Senna sophora Roxb.
  • Senna sousana H.S.Irwin & Barneby
  • Senna spectabilis (DC.) H.S.Irwin & Barneby
  • Senna spinescens (Hoffmanns. ex Vogel) H.S.Irwin & Barneby
  • Senna spiniflora (Burkart) H.S.Irwin & Barneby
  • Senna spinigera (Rizzini) H.S.Irwin & Barneby
  • Senna splendida (Vogel) H.S.Irwin & Barneby
  • Senna stenophylla (Britton) H.S.Irwin & Barneby
  • Senna stipulacea (Aiton) H.S.Irwin & Barneby
  • Senna stowardii (S.Moore) Randell
  • Senna stricta (Randell) Randell
  • Senna suarezensis (Capuron) Du Puy
  • Senna subtrijuga H.S.Irwin & Barneby
  • Senna subulata (Griseb.) H.S.Irwin & Barneby
  • Senna sulfurea (DC. ex Collad.) H.S.Irwin & Barneby
  • Senna surattensis (Burm.f.) H.S.Irwin & Barneby
  • Senna symonii (Randell) Randell
  • Senna talpana H.S.Irwin & Barneby
  • Senna tapajozensis (Ducke) H.S.Irwin & Barneby
  • Senna tenuifolia (Vogel) H.S.Irwin & Barneby
  • Senna timoriensis (DC.) H.S.Irwin & Barneby
  • Senna tocotana (Rose ex Britton & Killip) Silverst.
  • Senna tonduzii (Standl.) H.S.Irwin & Barneby
  • Senna tora (L.) Roxb. – sickle senna, foetid cassia
  • Senna trachypus (Mart. ex Benth.) H.S.Irwin & Barneby
  • Senna trianae H.S.Irwin & Barneby
  • Senna trolliiflora H.S.Irwin & Barneby
  • Senna tropica (Vell.) H.S.Irwin & Barneby
  • Senna truncata (Brenan) Lock
  • Senna tuhovalyana (Aké Assi) Lock
  • Senna uncata H.S.Irwin & Barneby
  • Senna undulata (Benth.) H.S.Irwin & Barneby
  • Senna uniflora (Mill.) H.S.Irwin & Barneby
  • Senna unijuga (Rose) H.S.Irwin & Barneby
  • Senna urmenetae (Phil.) H.S.Irwin & Barneby
  • Senna vargasii (Schery) H.S.Irwin & Barneby
  • Senna velutina (Vogel) H.S.Irwin & Barneby
  • Senna venusta (F.Muell.) Randell
  • Senna versicolor (Meyen ex Vogel) H.S.Irwin & Barneby
  • Senna viarum (Little) H.S.Irwin & Barneby
  • Senna viciifolia (Benth.) H.S.Irwin & Barneby
  • Senna viguierella (Ghesq.) Du Puy
  • Senna villosa (Mill.) H.S.Irwin & Barneby
  • Senna viminea (L.) H.S.Irwin & Barneby
  • Senna weddelliana H.S.Irwin & Barneby
  • Senna williamsii (Britton & Rose) H.S.Irwin & Barneby
  • Senna wislizeni (A.Gray) H.S.Irwin & Barneby
  • Senna wurdackii H.S.Irwin & Barneby – Wislizenus' senna, shrubby senna

Fossil record

A fossil seed pod of Senna sp. from the middle Eocene epoch has been described from the Rancho clay pit in Henry County, Tennessee (United States).


This article uses material from the Wikipedia article
 Metasyntactic variable, which is released under the 
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